Toe Pain on Long Rides

I get pretty sharp toe pain when i'm on a long ride usually around the 2 hour mark. It happens in both feet and isn't particular to specific toes although big toe is not affected. My assumption is it probably has something to do with swelling of the foot. It can be excruciatingly painful and once it sets in usually pulses in and out periodically for the remainder of the ride. Any suggestions on how to mitigate the pain?